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1 Downloaded from UvA-DARE, the institutional repository of the University of Amsterdam (UvA) File ID Filename Titlepage SOURCE (OR PART OF THE FOLLOWING SOURCE): Type Dissertation Title Evolutionary Ecology of the coral genus Madracis - an illustration of the nature of species in scleractinian corals Author M.J.A. Vermeij Faculty Faculty of Science Year 2002 Pages 255 ISBN FULL BIBLIOGRAPHIC DETAILS: Copyright It is not permitted to download or to forward/distribute the text or part of it without the consent of the author(s) and/or copyright holder(s), other then for strictly personal, individual use. UvA-DARE is a service provided by the library of the University of Amsterdam (

2 Evolutionary Ecology of the Coral Genus Madracis on Caribbean Reefs an illustration of the nature of species in scleractinian corals ACADEMISCH PROEFSCHRIFT ter verkrijging van de graad van doctor aan de Universiteit van Amsterdam op gezag van de Rector Magnificus prof. mr. P.F. van der Heijden ten overstaan van een door het college voor promoties ingestelde commissie in het openbaar te verdedigen in de Aula der Universiteit op woensdag 13 maart 2002 te uur door Mark Johannes Alphons Vermeij geboren te Gilze-Rijen Evolutionary Ecology of Madracis

3 Promotor: Prof. dr. R.P.M. Bak Leden promotiecommissie: Dr. R.W.M van Soest (Universiteit van Amsterdam) Prof. dr. F.R. Schram (Universiteit van Amsterdam) Dr. P.J. Mumby (University of Newcastle) Prof. dr. J. Videier (Rijksuniversiteit Groningen) Dr. J.A. Kaandorp (Universiteit van Amsterdam) Prof. dr. S.B.J. Menken (Universiteit van Amsterdam) Faculteit der Wiskunde, Natuurwetenschappen en Informatica Instituut voor Biodiversiteit en Ecosysteemdynamica (IBED) / Evolutiebiologie Onderzoekschool Biodiversiteit ISBN Niets uit deze uitgave mag worden vermenigvuldigd en/of openbaar gemaakt door middel van druk, microfilm, fotokopie of op welke andere wijze dan ook zonder voorafgaande schriftelijke toestemming van de auteur. No part of this book may be reproduced in any format by print, photocopy, microfilm or by any other means without the written permission from the author. Evolutionary Ecology oîmadracis

4 Evolutionary Ecology of Madracis I realize that I'm a fortunate person in doing the work that is a dream of many people. Working at the pristine reefs of Curaçao and Bonaire gave me the chance to get to know many aspects of coral reef life. But even at these places the ongoing worldwide decline of coral reefs is visible. Since reefs often occur in places where other problems prevail, the task of protecting them is not the easiest. I therefore dedicate this thesis to those that do this work under such circumstances. Secondly, there is a type of person that is willing to sacrifice a lot in pursuing their dream to work on coral reefs. I was lucky to have met a few of them and it is also to them that I dedicate this thesis. Without them reefs may disappear before anybody knows what they really are.

5 C,\RM.4Ä/ Vermeij, M.J.A. Evolutionary ecology of the coral genus Madracis on Caribbean reefs. PhD thesis, University of Amsterdam, the Netherlands. 255 pages. Copyright Mark J.A. Vermeij This thesis was prepared at the Institute for Biodiversity and Ecosystem Dynamics (IBED) of the University of Amsterdam (Mauritskade 61, 1090 GT Amsterdam, the Netherlands) and the department of Marine Ecology of the Netherlands Institute for Sea Research (NIOZ; P.O.Box 59, 1790 AB Den Burg, the Netherlands). The fieldwork was carried out at the Caribbean Institute for the Management and Research of Biodiversity, CARMABI (Piscaderabaai z/n, P.O. Box 2090, Willemstad, Curaçao, Netherlands Antilles) and the Bonaire Marine Park (BMP, Bonaire, Netherlands Antilles). The research was funded by the University of Amsterdam and NIOZ. Printed by Ponsen en Looijen BV Wageningen Evolutionary Ecology of Madracis 6

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